really confused and messed up shortcuts

That does not work. "Show autofill on field focus" is VERY annoying and frequently pops up in wrong places.
This is exactly my point - it's all very confusing and conflicting.
It seems I have to press the Fill in Browser shortcut first and if that does not do anything I then have to try the Open with shortcut in the browser extension. It used to be clean and simple - a single shortcut would either fill (if only one) or show a list if there where multiple possibilities.

This really is a mess and a step backwards!